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American Society of Animal Science
1111 N. Dunlap Ave., Savoy, IL 61874
Phone: 
+1 217.356.3182
Fax: +1.217.398.4119
www.asas.org
jas.fass.org
asas@assochq.org

Journal of Animal Science 

Serving Human Needs Through Animal Agriculture

Purpose and Scope:

The Journal of Animal Science (JAS) is among the top-ranked journals of animal science research in the world. The official journal of the American Society of Animal Science, JAS publishes results of original research in Genetics, Growth and Physiology, Nutrition, Production, Products, and Special Topics. JAS consistently ranks in the top tier in the category of Agriculture, Dairy, and Animal Sciences titles by impact factor for the ISI rankings and is in the top 1% of STM publishing (50,000+ titles) by total ISI citations. JAS is now available to all institutional subscribers online on the HighWire Press platform, allowing a feature-rich environment for users and management and tracking tools for institutions.

Intended Readership:

Readers of JAS  represent education, industry, and government, including research, teaching, administration, veterinary medicine, extension, management, quality assurance, product development, and technical services. JAS is of interest to those in biochemistry, breeding, economics, engineering, environment, food science, genetics, microbiology, nutrition, pathology, physiology, processing, public health, quality assurance, and sanitation. 

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Bibliographic Information:

Print ISSN: 0021-8812; Online ISSN: 1525-3163
2008 Volume 86
Issues per year: 12 + 2 supplements (Annual Meeting and Sectional Abstracts)
Frequency: monthly 
Print Subscription Format: 8 1/2” × 11”; 300 line screen photographs; enamel, acid-free paper; includes access to online version
JAS is peer reviewed

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Indexing and Abstracting:

JAS is indexed and/or abstracted in Agricola • Biological Abstracts • Biological and Agricultural Index • BIOSIS Database • CABI Abstracts • Current Contents • HighWire Press portal • Infotrieve • ISI Web of Science • Proquest • PubMed/Medline

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Advance Pricing:

2008 Yearly Subscription Rate—Print (includes online access)
U.S.A., Canada, and Mexico: $525
Elsewhere: $550

(Prices include surface mail postage.)
$450 online only

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Editorial Board:

Editor-in-Chief
L. P. Reynolds
North Dakota State University
Department of Animal and Range Sciences
Fargo, ND 58105-5727
Phone: (701) 231-7646
Fax: (701) 231-7590
Larry.Reynolds@ndsu.edu

Animal Genetics:
Molecular Genetics
Quantitative Genetics
Division Editor
M. E. Davis (2009)
The Ohio State University
2027 Coffey Road
Columbus, OH 43210
Phone: (614) 292-4984
Fax: (614) 292-7116
davis.28@osu.edu

Associate Editor
S. A. Barwick (2009)
AGBU
University of New England
Armidale, New South Wales 2351, Australia
Phone: 61-2-6773-3481
Fax: 61-2-6773-3266
steve.barwick@dpi.nsw.gov.au

Associate Editor
E. Casas (2009)
USDA, ARS, US MARC
PO Box 166
State Spur 18D
Clay Center, NE 68933-0166
Phone: (402) 762-4168
Fax: (402) 762-4173
Eduardo.casas@ars.usda.gov

Associate Editor
D. H. Crews Jr. (2009)
Lethbridge Research Centre
Agriculture and AgriFood Canada
5403 1st Avenue South
Lethbridge, Alberta T1J 4B1, Canada
Phone: 403-317-2288
Fax: 403-382-3156
dcrews@agr.gc.ca

Associate Editor
B. Kirkpatrick (2009)
University of Wisconsin-Madison
Animal Science Bldg.
1675 Observatory Drive
Madison, WI 53706
Phone: 608-263-4323
Fax: 608-262-5157
bwkirkpa@wisc.edu

Associate Editor
G. Rosa (2009)
Dairy Science
University of Wisconsin-Madison
456 Animal Sciences Building
1675 Observatory Dr.
Madison, WI 53706
Phone: (608) 265-8617
Fax: (608) 263-3308
grosa@wisc.edu

Animal Growth, Physiology, and Reproduction:
Growth and Developmental Biology
Physiology, Endocrinology, and Reproduction

Divison Editor
S. A. Zinn (2008)
University of Connecticut
Department of Animal Science
3636 Horsebarn Rd.
Ext. Unit 4040
Storrs, CT 06269-4040
Phone: (860) 486-0861
Fax: (860) 486-4375
szinn@canr.cag.uconn.edu

Associate Editor
M. J. Azain (2008)
University of Georgia
Rhodes Center for Animal and Dairy Science
Athens, GA 30602-2771
Phone: (706) 542-0963
Fax: (706) 542-0399
mazain@uga.edu

Associate Editor
D. L. Thompson Jr. (2008)
Louisiana State University
Department of Animal Science
Baton Rouge, LA 70803-4210
Phone: (225) 578-3445
Fax: (225) 578-3279
dthompson@agctr.lsu.edu

Associate Editor
C. R. Barb (2009)
Russell Research Center, Animal Physiology Research Unit
USDA/ARS
PO Box 5677
Athens, GA 30604-5677
Phone: (706) 583-8276
Fax: (706) 542-0399
Richard.barb@ars.usda.gov

Animal Nutrition:
Nonruminant Nutrition
Ruminant Nutrition
Division Editor
P. S. Miller (2008)
University of Nebraska
C206g Animal Sciences
Lincoln, NE 68583
Phone: (402) 472-6421
Fax: (402) 472-6362
pmiller1@unl.edu

Division Editor
E. C. Titgemeyer (2008)
Kansas State University
Department of Animal Science
132 Call Hall
Manhattan, KS 66506-1600
Phone: (785) 532-1220
Fax: (785) 532-5681
etitgeme@oznet.ksu.edu

Associate Editor
M. Bauer (2008)
North Dakota State University
Department of Animal and Range Sciences
155 Hultz Hall
Fargo, ND 58105
Phone: (701) 231-7691
Fax: (701) 231-7596
mbauer@ndsuext.nodak.edu

Associate Editor
K. Beauchemin
Agriculture and Agri-Food Canada
Research Center Box 3000
Lethbridge, AB T1J 4B1
Canada
Phone: (403) 317-2235
Fax: (403) 382-3156
beauchemin@agr.gc.ca

Associate Editor
L. I. Chiba (2008)
Auburn University
Department of Animal Sciences
303C Ann S. Upchurch Hall
Auburn University, AL 36849-5415
Phone: (334) 844-1560
Fax: (334) 844-1519
chibale@auburn.edu

Associate Editor
J. Eisemann (2008)
North Carolina State University
Department of Animal Science
Box 7621
Raleigh, NC 27695
Phone: (919) 515-4044
Fax: (919) 515-7780
joan_eisemann@ncsu.edu

Associate Editor
B. Kerr (2008)
USDA-ARS-SOMMRU
Nat'l Swine Res/Info Ctr
2150 Pammel Drive, Room 2167
Ames, IA 50011
Phone: (515) 294-0224
Fax: (515) 294-1209
kerr@nsric.ars.usda.gov

Associate Editor
M. Kerley (2008)
University of Missouri
920 E. Campus Drive
Lab 111, ASRC
Columbia, MO 65211-5300
Phone: (573) 882-0834
Fax: (573) 884-7712
kerleym@missouri.edu

Associate Editor
D. Bohnert (2009)
Eastern Oregon Agricultural Research Center
Oregon State University
67826-A Hwy 205
Burns, OR 97720
Phone: 541-573-8910
Fax: 541-573-3042
dave.bohnert@oregonstate.edu

Animal Production:
Behavior
Environment
Feedstuff Evaluation
Health and Well-Being
Management
Pharmacology and Toxicology
Rangeland, Pasture, and Forage Utilization
Division Editor
D. M. Hallford (2008)
New Mexico State University
Department of Animal and Range Sciences
Box 3I
Las Cruces, NM 88003
Phone: (505) 646-1004
Fax: (505) 646-5441
dhallfor@nmsu.edu

Associate Editor
S. Eicher-Pruiett (2008)
USDA-ARS Livest. Behav. Res.
216 Poultry
West Lafayette, IN 47907
Phone: (765) 496-3665
Fax: (765) 496-1993
spruiett@purdue.edu

Associate Editor
S. A. Gunter (2008)
University of Arkansas
Southwest Research/Ext. Center
362 Highway 174 North
Hope, AR 71801
Phone: (870) 777-9702
Fax: (870) 777-0963
sgunter@uaex.edu

Associate Editor
G. P. Lardy (2008)
North Dakota State University
Animal and Range Sciences Dept
100 E. Hultz Hall
Fargo, ND 58105
Phone: (701) 231-7660
Fax: (701) 231-7590
glardy@ndsuext.nodak.edu

Associate Editor
M. L. Looper (2008)
USDA, ARS, SPA
6883 S. State Hwy 23
Booneville, AR 72927
Phone: (479) 675-3834
Fax: (479) 675-2940
mlooper@spa.ars.usda.gov

Associate Editor
J. B. Taylor (2008)
USDA - Agricultrual Research Service
HC 62 Box 1954
Dubois, ID 83423
Phone: (208) 374-5306
Fax: (208) 374-5582
btaylor@pw.ars.usda.gov

Animal Products:
Meat Science and Muscle Biology
Pre- and Postharvest Product Safety

Division Editor
D. E. Gerrard (2008)
Purdue University
202A Smith Hall
West Lafayette, IN 47907
Phone: (765) 494-8280
Fax: (765) 494-6816
dgerrard@purdue.edu

Associate Editor
B. Bowker (2008)
USDA-ARS
10300 Baltimore Avenue
Bldg 201 BARC-EAST
Beltsville, MD 20705
Phone: (301) 504-5626
Fax: (301) 504-8438
bbowker@anri.barc.usda.gov

Associate Editor
M. Du (2009)
Animal Science
University of Wyoming
1000 E. University Avenue
Laramie, WY 82071
Phone: 307-766-3429
Fax: 307-766-2355
mindu@uwyo.edu

Associate Editor
K. W. McMillin (2009)
Animal Sciences
Louisiana State University AgCenter
Francioni Hall So. Campus Dr.
Baton Rouge, LA 70803-4210
Phone: (225) 578-3438
Fax: (225) 578-3279
KMcmillin@agcenter.lsu.edu

Special Topics:
Biographical/Historical Sketches
Contemporary Issues
Teaching

Division Editor
R. Dailey (2008)
West Virginia University
Animal & Veterinary Science
G040 Agricultural Sciences Building
PO Box 6108
Morgantown, WV 26506
Phone: (304) 293-2395
Fax: (304) 293-2232
rdailey@wvu.edu

Symposia
M. Mirando (2008)
USDA-CSREES-NRI
Stop 2241
1400 Independence Ave SW
Washington, DC 20250-2241
Phone: (202) 401-4336
Fax: (202) 401-3641
mmirando@csrees.usda.gov

Editorial Board
Sam Baidoo, University of Minnesota
Brian Bequette, University of Maryland
Jim Berardinelli, Montana State University
Chris Calvert, University of California, Davis
Nuria Canibe, University of Aarhus, Denmark
Fernando Cardoso, Embrapa, Bage, Brazil
Jeffery Clapper, South Dakota State University
Wayne Coblentz, US Dairy Forage Research Center, Marshfield, WI
John Cole, USDA-ARS, Beltsville, MD
Candace Croney, Oregon State University
Shawn Donkin, Purdue University
Sandra Edwards, Newcastle University
Steve Ellis, Clemson University
Mark Enns, Colorado State University
Galen Ericksen, University of Nebraska
George Fahey, University of Illinois
Eugene Felton, West Virginia University
Joe Ford, USDA-ARS, Clay Center, NE
Joe Garner, Purdue University
Thierry Gidenne, INRA, Toulouse, France
Kristen Govoni, Loma Linda University
Elaine Grings, USDA-ARS, Miles City, MT
Kevin Halpin, International Ingredient Corp., Fenton (St. Louis), MO
Kathryn Hanford, University of Nebraska-Lincoln
Matt Hersom, University of Florida
Rhonda Hoffman, Middle Tennesse State University
Elisabeth Huff-Lonergan, Iowa State University
Walt Hurley, University of Illinois
Brad Johnson, Kansas State University
Sally Johnson, University of Florida
Stephen Kachman, University of Nebraska-Lincoln
Bas Kemp, Wageningen University
James Kijas, CSIRO Livestock Industries, St Lucia, Australia
John Killefer, University of Illinois
Elizabeth Koutsos, Mizuri Exotic Animal Nutrition, California
Scott Lake, Purdue University
Laurie Lawrence, University of Kentucky
Kinchon Lee, Ohio State University
Tim Leeds, US Sheep Experiment Station. Dubois, ID
Xingen Lei, Cornell University
Clay Lents, University of Georgia
Steven Lonergan, Iowa State University
Jim MacDonald, Texas Agric Exp Sta., Amarillo
Jeremy Marchant Forde, USDA Livestock Behavior Research Unit, West Lafayette, IN
Tim McAllister, Agriculture and Agri-Food Canada
Tara McDaneld, USDA-ARS, Clay Center, NE
Joy Mench, University of California, Davis
Brian Nielsen, Michigan State University
Donald Joshua Nkrumah, Univeristy of Alberta
Martin Nyachoti, University of Manitoba
Anita Oberbauer, University of California, Davis
Erasmus Okine, University of Alberta
K. C. Olson, Kansas State University
Isabelle Oswald, INRA, Toulouse, France
David Patterson, University of Missouri
Robert L. Payne, Degussa Corporation
Miguel Perez-Enciso, Universitat Autonoma de Barcelona
Richard Reynnells, USDA-CSREES-PAS, Laurel, MD
Mike Rinker, DPI GLOBAL, Berea, KY
Gary Rohrer, USDA-ARS, Clay Center, NE
Roberto Sainz, University of California, Davis
Janeen Salak-Johnson, University of Illinois
Brian L. Sayre, Virginia State University, Petersburg
Keith Schillo, University of Kentucky
Robert Schnabel, University of Missouri
Patricia Schoknecht, Wagner College, Staten Island, NY
Eric Scholljegerdes, USDA-ARS, Mandan, ND
Marcia Shannon, University of Missouri
Paul Siciliano, North Carolina State University
Kathy Soder, USDA-ARS
Sergio Soto-Navarro, New Mexico State University
Matthew Spangler, University of Tennessee at Martin
Michael Spurlock, Iowa State University
Hans Stein, University of Illinois
Kelly Swanson, University of Illinois
Kendall Swanson, University of Guelph
Jeffrey Vallet, USDA-ARS, Clay Center, NE
Eric van Heugten, North Carolina State University
Eric VanZant, University of Kentucky
Mike Wade, Universidad Nacional del Centro de la Provincia de Buenos Aires
Zhiquan Wang, University of Alberta
Tryon Wickersham, Texas A&M University
Stephan Wildeus, Virginia State University, Petersburg
Scott Wisnant, North Carolina State University
Ruurd Zijlstra, University of Alberta

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Published by:

The Journal of Animal Science (JAS) is published by the American Society of Animal Science. Established in 1908, the American Society of Animal Science (ASAS) is a professional organization for animal scientists designed to help members provide effective leadership through research, extension, teaching and service for the dynamic and rapidly changing livestock and meat industries. The mission of ASAS is to discover, disseminate, and apply knowledge for sustainable use of animals for food and other human needs.

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